Rugosa / 30th November 2016

BADPA M., POTY E., ASHOURI A., KHAKSAR K. 2016. Fasciculate Kleopatrinid corals from the Bashkirian (Late Carboniferous) of Sardar Formation (Ozbak-Kuh Mountains, East-Central Iran). Revista brasileira de paleontologia 19, 2: 151-166.
[keywords: Iran, Ozbak-kuh, Sardar, Bashkirian, rugose corals]
In the East-Central Iran, the Sardar Formation (upper Visean-Moscovian) consists of shallow-water limestone with intercalations of shale containing rugose corals, tabulate corals and brachiopods. Ten sections were sampled in the Ozbak-kuh Mountains, from north to south. Among the rugose corals, an assemblage of fasciculate Kleopatrinidae has been collected. The latter contains the species: Paraheritschioides antoni antoni, P. antoni minor, P. gracilis and two new species for the genera Fomichevella and Heintzella. Heintzella is described from Iran for the first time. However, its age, as determined by conodonts and foraminifers, is early to middle Bashkirian (early late Carboniferous). The most similar, time-equivalent faunal associations are that of the Ellesmere Island, Sverdrup Basin in Arctic Canada, Alexander terrane and Brooks Range in southeastern Alaska and eastern Klamath terrane in northern California, where similar tropical warm water conditions have been identified during the Bashkirian in the northern hemisphere. During these times central Iran block and Northern provinces, [were] characterized by a dominant carbonate facies and more diversified colonial coral faunas. [original abstract]

BERKOWSKI B., ZAPALSKI M. K., WRZOLEK T. 2016. New Famennian colonial coral (Rugosa) from the Holy Cross Mountains (Poland): an example of local evolution after Frasnian-Famennian extinction. Naturwissenschaften 2016 Apr;103 (3-4): 33. doi: 10.1007/s00114-016-1356-1. Epub 2016 Mar 16.
[keywords: Colonial Rugosa; Devonian extinction; Famennian; Recovery; Taxonomy]
Colonial rugose corals are extremely rare in the fossil record after the Late Devonian (Frasnian-Famennian) extinction event. Here, we report a new genus and species, Famastraea catenata, from the late Famennian of the western part of the Holy Cross Mountains (Kowala) in Poland. Although this taxon is colonial, it displays many morphological characters very close to the typically late Famennian solitary species Palaeosmilia aquisgranensis (Frech, 1885), described earlier from the same locality. Hence, we postulate that F. catenata is derived from P. aquisgranensis. In contrast to other Famennian colonial rugose corals, the new taxon represents an example of local evolution within the group of so-called 'Strunian' corals. Consequently, we postulate that the new taxon represents a new colonial rugose fauna, which, however, did not survive the subsequent Late Devonian crisis (i.e. Hangenberg event). F. catenata most probably inhabited deeper water settings, possibly near the boundary between the euphotic and dysphotic zones, as inferred from many other benthic taxa described from this locality. [original abstract; Wrzolek]

COEN-AUBERT M. 2016. Potyphyllum, a new phillipsastreid genus of rugose corals in the Upper Frasnian of Belgium with precisions about the age of the Petit-Mont Member. Geologica Belgica 19, 1/2: xxx-xxx [9pp + 2 Pls].
[keywords: Rugosa, taxonomy, lithostratigraphy, biostratigraphy, Frasnian]
The new genus Potyphyllum is erected with Cyathophyllum ananas Goldfuss, 1826 as type species, for which a neotype is selected here from the north side of the Namur Basin. It is a massive pseudocerioid rugose coral like Frechastraea Scrutton, 1968 with rather large corallites, spindle-shaped dilated septa as well as some horseshoe and inner dissepiments. In the Belgian Upper Frasnian, Potyphyllum ananas and P. veserense (Coen-Aubert, 1974) occur mainly in the Upper Palmatolepis rhenana conodont Zone. The latter two coral taxa, together with diverse species of Frechastraea and Phillipsastrea D'Orbigny, 1849, permit dating of the red marble lenses from the Petit-Mont Member which are developed at different levels of the Champ Broquet Formation, in the Dinant Synclinorium. In this lithostratigraphic unit are also incorporated the Neuville and Les Valisettes Members, formerly considered as two separate formations. At the present time, Potyphyllum is only known from Western and Eastern Europe. [original abstract; Wrzolek]

DENAYER J. 2015. Taxonomy, Biostratigraphy and Palaeobiogeography of the Late Tournaisian rugose corals of north-western Turkey. Palaontologische Zeitschrift 89, 3: 313-333.
[keywords: Corals; Carboniferous; Tournaisian; Ivorian; palaeogeography; stratigraphy]
This article provides the first taxonomic description of Ivorian (Late Tournaisian, Early Carboniferous) rugose coral associations from north-western Turkey (Zonguldak and Bartin). Eleven species belonging to ten genera are described, one species is new. Three biostratigraphic assemblages are recognized. The oldest assemblage includes Cyathaxonia cornu, Cyathoclisia uralensis, 'Lophophyllum' konincki and Uralinia multiplex. This corresponds to the RC3 Biozone (early Ivorian). The middle assemblage in characterised by Amplexus coralloides, Sychnoelasma hawbankense and Zaphriphyllum daleki sp. nov. and is correlated with the early late Ivorian RC4α Biozone. The youngest assemblage (RC4β1 Biozone, latest Ivorian) is composed of Corphalia fourmarieri, Corphalia sp. and Amydgalophyllum? sp. These three assemblages have a low specific and generic diversity compared to time-equivalent assemblages but contain genera with a wide distribution in the Palaeotethys Ocean, such as Cyathoclisia, Uralinia and - in a lesser extent - Zaphriphyllum, as well as typically European taxa such as Corphalia and Sychnoelasma. These latter two, identified for the first time outside of Europe, allow associating north-western Turkey with the European Coral Province. [original abstract; Wrzolek]

DENAYER J. 2015. Rugose corals at the Tournaisian-Viséan transition in the Central Taurides (S Turkey) - Palaeobiogeography and palaeoceanography of the Gondwana margin. Journal of Asian Earth Sciences 98: 371-398.
This first taxonomic description of the Upper Tournaisian-Lower Viséan rugose coral fauna of the Yaricak Formation (Aladag Unit, Central Taurides, South Turkey) is here provided. The corals are distributed into four stratigraphical assemblages. The Upper Tournaisian assemblages include widely distributed taxa (Uralinia, Caninia, Proheterelasma, Zaphrentites). The assemblage crossing the Tournaisian-Viséan boundary is characterized by both Eurasian and cosmopolitan and widely distributed taxa (Calmiussiphyllum, Siphonophyllia, Bifossularia Amygdalophyllum, Caninophyllum, Keyserlingophyllum) and Asian taxa (Kueichouphyllum). The youngest assemblage (Lower Viséan) is dominated by Eokoninckocarinia gemmina sp. nov. These assemblages are typical of the South Palaeotethys so-called 'Kueichouphyllum Zone' extending along the Asian margin of Gondwana (Cimmerian Terrane) during Lower Carboniferous times. As in the other Cimmerian blocks, colonial taxa are absent. [original abstract; Denayer]

DENAYER J., OGAR V. 2016. Vassiljukia, a new colonial rugose coral from the early Viséan (Mississippian) of the Donets Basin (Ukraine) and NW Turkey. Comptes rendus Paléovol. 15, 8: 911-917.
The cerioid colonial coral previously described as Lithostrotion columnariformis Vassiljuk, 1960 (Early Viséan, Donets Basin, Ukraine) is re-attributed to the new genus Vassiljukia. This genus is introduced for colonial amygdalophylloid developing a stable cerioid habitus differing from amygdalophylloid proto-colonies by the ability to produce a second generation of offsets. The origin and affinity of Vassiljukia columnariformis within the Amygdalophyllidae are also discussed. [original abstract; Denayer]

DENAYER J., WEBB G. E. 2015. Cionodendron and related lithostrotionid genera from the Mississippian of eastern Australia: systematic, stratigraphy and evolution. Alcheringa 39: 1-62.
This paper is a systematic revision of the Australian Mississippian colonial rugose corals previously described as Lithostrotion, Siphonodendron and Orionastraea. The systematic revision highlighted their homeomorphy with the Eurasian genera Siphonodendron, Lithostrotion and Heterostrotion resulting from parallel evolution within the Lithostrotionidae. Fasciculate species are reassigned to the genus Cionodendron as they share a robust columella, septotheca and two series of tabellae. The massive species are classified in the newly created genus Australastraea that likely evolved from Cionodendron. Australastraea is characterized by a septotheca that is commonly discontinuous, conferring a pseudo-cerioid or astreoid habit. Small-sized fasciculate species previously referred to Lithostrotion williamsi are here reassigned to the new genus Pickettodendron, which differs from Cionodendron by the lack of minor septa and presence of a complete tabularium. A polyphyletic origin of the newly introduced Cionodendroninae subfamily is considered, with two species of Amygdalophyllum possibly being at the origin of Cionodendron - Australastraea lineage and of Pickettodendron. Cionodendroninae assemblages of eastern Australia are strongly endemic and could represent one of the scarce remains of the Panthalassa Province. [original abstract; Denayer]

FEDOROWSKI J. 2016. On the genus Bothrophyllum Trautschold, 1879 (Anthozoa, Rugosa). Acta Geologica Polonica 66, 1: 1-41.
[keywords: Carboniferous Bothrophyllum; Morphological frames; Origin; Distribution; Relationships]
The rugose coral genus Bothrophyllum Trautschold, 1879 is revised on the basis of data from the literature and the author's personal investigation of both topotypes of its type species B. conicum and related and/or similar taxa from other areas. The intraspecific variability of the type species, its neotype, the intra-generic framework and a new generic diagnosis are established. Many more than 100 taxa related and/or similar to Bothrophyllum were analyzed and the most important of them are discussed. Detailed analysis of the type species based on the neotype and supported by additional topotype specimens illustrated here, allows restriction of both the type species and the genus, and leads to the proposition that Bothrophyllum - like taxa with a shortened cardinal septum should be considered of subgeneric (not named) status. Detailed analysis of the specimens and species described and illustrated from the type site (Myachkovo Quarry, Moscow Basin) form the basis for further considerations. On the basis of that analysis and characters established for the type species, taxa from all other European, African, Asiatic and North American areas either named Bothrophyllum or bearing characters of that genus were analyzed. The supposed origin and discussion of the relationships conclude the paper. A list of synonyms and exclusions from Bothrophyllum and lists of species included, excluded, or possibly belonging to Bothrophyllum and Bothrophyllum - like corals with a shortened cardinal septum are presented. [original abstract; Wrzolek]

FEDOROWSKI J. 2017 (accepted). Early Bashkirian Rugosa (Anthozoa) from the Donets Basin (Ukraine). Part 5. The Family Bothrophyllidae Fomichev, 1953. Acta Geologica Polonica. [accepted for the second issue of 2017 volume of AGP]
Four genera (one new, one questionable), seven named species (five new) and five taxa left in the open nomenclature are described from the uppermost Serpukhovian (Limestone D5/3) to the middle Bashkirian (Limestone G1/1) strata. A new genus: Nina and new species: Bothrophyllum kalmiussi, B. gorbachevensis, Nina donetsiana, N. dibimitaria and N. magna are introduced. The genus Cystilophophyllum Fomichev, 1953 is redefined and transferred to the Family Bothrophyllidae. Three general topics discussed in the Considerations are: 1. The taxonomic value of the microstructure of septa and the necessity for distinction between that character and the diagenetic alterations. 2. Depositional conditions under which the rugose coral skeletons were deposited. 3. The similarity of skeletal constructions vs relationships in some taxa of the Subfamily Dibunophyllinae Wang, 1950 and the Family Bothrophyllidae Fomichev, 1953. The lack of final conclusions in the matter of the relationships is due to the incompleteness and restricted number of specimens studied here and the incomplete investigations of a great majority of the earlier described bothrophyllid taxa. [original abstract; Fedorowski]

KOSSOVAYA O. L., NOVAK M., WEYER D. 2016. Large-sized Early Permian "caninioid" corals from the Karavanke Mountains, Slovenia. Journal of Paleontology, online since 19 September 2016, pp. 1-19; DOI:
A new monospecific "caninioid" genus, Preisingerella n. gen., from the lower Permian of the Karavanke Mountains (Southern Alps, Slovenia) is erected. The type species is Preisingerella stegovnikensis n. sp. The new taxon can be differentiated from other Caninia-type genera by its specific ontogeny and features of its dissepimentarium. Corals with such morphology had a wide distribution during the Carboniferous and early Permian, occurring in mostly shallow-water carbonate rocks. The phylogenetic relationships within this group are mostly unclear due to similarities in the adult stages. The earlier stages reveal the main distinguishing features that are decisive for a generic assignment, but these have rarely been well preserved and properly considered. The new taxon is compared with related genera of the Cyathopsidae and species of Caninella Gorskiy, 1938 characterized by lateral dissepiments. Large numbers of specimens of the new species, representing a monospecific assemblage, have been collected from the Born Formation at Mt. Stegovnik. Sedimentological and microfacies characteristics, as well as macro- and microfossil assemblages, underline this correlation. The fusulinoidean assemblage of the Born Formation, with Sphaeroschwagerina carniolica (Kahler and Kahler, 1937), as the predominant species, corresponds to the time span between the Sphaeroschwagerina moelleri - Schwagerina fecunda and Pseudofusulina moelleri zones, indicating a late Asselian to early Sakmarian age in the Southern Urals. [original abstract]

OGAR V. V. 2016. New Rugose corals and refinements of the Tournaisian biostratigraphy of the Donets Basin (Ukraine). Geologica Belgica 19, 1-2: 21-28.
[keywords: Carboniferous, Mississippian, Dinantian, Rugose corals, biostratigraphy, palaeobiogeography]
Specimens collected by Vassilyuk from the lower part of the Tournaisian strata of the southern Donets Basin and described as Campophyllum caninoides Sibly are re-examined. These specimens [are] re-attributed to the Conilophyllum priscum (Münster, 1840), which is a well-known species in the lower part of the Lower Tournaisian of Western and Eastern Europe, Kazakhstan, North America and China (Junggar). Corphalia simplex (Perna, 1923) is for the first time identified at the top of Tournaisian in the Donets Basin. This species was described at similar stratigraphic levels on the eastern slope of the Southern Urals and in Belgium. The application of described species for interregional correlation of the Tournaisian subdivisions is discussed herein. Evolutionary relations between Corphalia and Dorlodotia near the Tournasian-Visean boundary have been confirmed indirectly. [original abstract; Wrzolek]

RODRIGUEZ S., SOMERVILLE I. D. 2015. The Axophyllinae from SW Spain: a review. Boletín de la Real Sociedad Espanola de Historia Natural 108: 81-138.
A revision of the state of knowledge of the solitary axophyllids that have been grouped in the subfamily Axophyllinae (family Axophyllidae) and the description of the specimens collected in Sierra Morena (south-western Spain) is accomplished. This group of rugose corals has been reported frequently in the Carboniferous from Australia to North America and from China to Western Europe. However, some of the supposed records belong to other coral families that show some features in common, such as Geyerophyllidae and Aulophyllidae. After a detailed revision of the bibliographic references and some of the types, the genera included in this subfamily are: Axophyllum, Gangamophyllum, Pareynia, Semenophyllum, Protocarcinophyllum and the new genus Morenaphyllum. The genus Axoclisia remains as a doubtful member of the family. Axophyllinae are common in south-western Spain. They have been recorded and cited from the Guadiato Area and Los Santos de Maimona Basin comprising upper Viséan (Asbian, Brigantian) and Serpukhovian rocks, but most of those records remain undescribed. The record of Axophyllinae in Sierra Morena comprises ten species belonging to Axophyllum (four of them are new; A. cozari, A. julianaense, A. spinosum, A. spiralum), two species belonging to Gangamophyllum, one belonging to Pareynia (a new one, P. viacrucense) and two belonging to the new genus Morenaphyllum (M. antolinense, M. boyerense). [original abstract; Rodriguez]

WEYER D. 2016. Review of some Frasnian ahermatypic coral localities from Germany and description of a new genus Spinaxon (Anthozoa, Rugosa, Upper Devonian). Geologica Belgica 19, 1-2: xxx-xxx [13pp + 4 Pls].
[keywords: Frasnian, Kellwasser, ahermatypic Rugosa, Thuringia, Montagne Noire]
One of the greatest gaps in the present worldwide knowledge of Rugosa history are the unstudied ahermatypic coral faunas of Frasnian (and Upper Givetian) times, though they exist in many places. Most occurrences in Germany are briefly reviewed (Variscan Rhenish, Harz, Thuringian and Franconian Mountains; Baltoscandian shelf deposits in deep boreholes on the Baltic Sea island of Rügen). Intensive collecting in Thuringia presented rich Upper Frasnian samples from two localities (former Geipel quarry in the town of Schleiz, Vogelsberg-quarry near the village Tegau), dominated by Metriophyllum Milne-Edwards & Haime, 1850, Neaxon Kullmann, 1965, and the newly established genus Spinaxon (suborder Cyathaxoniina, family Petraiidae), with the monotypic type species Spinaxon potyi sp. nov. Cephalopod limestones yielded >100 specimens of this taxon; one specimen was found in the Montagne Noire (Frasnian Famennian boundary stratotype section Coumiac). The taxon descends from Neaxon Kullmann, 1965 or its nearest relative Petraia Münster, 1839; it is characterized by unique hypertrophic long trabicular spines on the margins of major septa within an aulos of circulotheca type. [original abstract; Wrzolek]

WRIGHT A. J., PLUSQUELLEC Y., GOURVENNEC R. 2016. Devonian operculate corals (Calceolidae, Cnidaria) from the Massif Armoricain, France. Alcheringa 40: xxx-xxx. ISSN 0311-5518
[keywords: Gerviphyllum gen. nov, Rhizophyllum, Chakeola, Calceola, Calceolidae, operculum, Devonian, Massif Armoricain, Rade de Brest, Nehou, France]
The operculate coral Calceola gervillei Bayle, 1878 is described for the first time on the basis of the type material from the Cotentin region of Normandy (North Armorican Domain), from Early Devonian (likely upper Lochkovian to lower Pragian) strata, and is chosen as the type species of the monotypic new genus Gerviphyllum. The new genus is also present in the l'Armorique Formation (lower Pragian) of the Plougastel Peninsula (Central Armorican Domain) as Gerviphyllum sp. cf. G. gervillei. One locality in the upper Emsian (Polygnathus serotinus Conodont Zone) Le Fret Formation, on the northern coast of the Crozon Peninsula, has yielded operculate coral specimens described here as ?Chakeola sp., the first (tentative) record of the genus outside eastern Australia, south China and Vietnam. The operculate coral Calceola collini sp. nov. is described from six localities in the early Middle Devonian (Eifelian: Polygnathus costatus Conodont Zone) Saint-Fiacre Formation of the Plougastel and Crozon Peninsulas (Central Armorican Domain), despite the fact that knowledge of the internal characters, especially of the operculum, of the type species Calceola sandalina is very limited. From an extensive review of published references to Calceola from France, we conclude that only the record of Collin (1929) is valid. [original abstract; Wrzolek]